I. Evolution is the most important concept in all of biology. None of biology makes sense except in the context of evolution. Just why is evolution so important to biologists? A few examples'
A. Taxonomy: Evolution makes sense of the similarities and differences and provides a rational explanation for the distribution of taxa. It also supplies explanations for the sometimes wild exceptions we see (eg cacti and euphorbs, which look so similar but are clearly only very distantly related to each other. The explanation is convergent evolution).
B. Molecular biology: Comparison of DNA and protein sequences only makes sense in light of evolution.
C. Medicine: Our understanding of infectious organisms and our tactics to combat them are entirely based upon understanding of evolution.
II. No idea comes from thin air. The historical perspective helps to understand where the concepts behind evolutionary theory came from. Here are some of the basic historical issues.
A. The typical view of the nature of species at the beginning of the nineteenth century is often described as the Doctrine of Fixed Species. This concept had its primary roots in Greek philosophy: Aristotle's Scala Naturae (The 'Great Chain of Being') and Plato's essentialism.
B. During the eighteenth century, Carolus Linnaeus (1707-1778) developed the basis for our nested, hierarchical classification system, or Taxonomy. This system is also a 'natural' system, meaning that, regardless of the characters of organisms chosen as the basis for the classification, species almost always end up classified in the same pattern, an oddity which didn't escape early 19th century naturalists, and which led to all kinds of evolutionary thinking.
C. The geologic discipline of stratigraphy was developed during the late eighteenth and early nineteenth centuries. Though this accomplishment was shared by many geologists throughout Europe, the name most commonly associated with developing the concept of stratigraphy is William Smith (1769-1839), who was an engineer engaged in building canals all over England.
D. George Cuvier (eventually Baron George Cuvier) (1769-1832) was a scientist of the late 18th and early 19th centuries. His specialty was vertebrate anatomy. He developed the important technique of comparative anatomy, which is based upon the understanding that the parts of an animal aren't just unrelated bits stuck together'they reflect the life style and type of the animal, and are thus all related to each other. Because of his expertise in anatomy, Cuvier was the obvious person to ask to examine strange remains found during reconstruction after a major Paris fire. These remains turned out to belong to extinct organisms, and led to the development of the field of study we call Paleontology. Cuvier was the first to reconstruct dinosaurs. Cuvier is also remembered for coming up with the concept of Catastrophism, as an attempt to reconcile his religious beliefs with the evidence he discovered as a scientist.
E. Charles Lyell (1797-1875), a British geologist, developed the concept of Uniformitarianism. This theory of geology is often summarized by the catch phrase, 'The present is the key to the past.' The meaning of this phrase is that, if you can understand how geological features are being shaped and formed today, you can look at the formations from a long time ago and assume that they were created in the same way. In other words, the forces acting upon and shaping the Earth today are the same as the forces that acted upon and shaped the Earth in the past. Uniformitarianism is a central tenet of modern geology.
III. Charles Darwin (1809-1882) and the Origin of Species
A. Who was Charles Darwin?
1. Darwin was born in the first century of the nineteenth century to a wealthy family. He began an education in medicine (the 'family trade') but actually completed his degree in theology. His hobby was natural history. He collected beetles, went on geological field trips and became very friendly with the natural history faculty at his universities.
2. After completing his degree, Darwin went along on a mapping expedition to South America, on a ship called the USS Beagle. During the five-year trip, he collected tons (literally) of specimens of animals, plants and fossils, which he shipped back to England for later analysis.
3. Upon returning to England, he officially became a naturalist. Almost all of his specimens were sent to experts for analysis. He kept the barnacles for himself, setting out to become an expert in barnacles. It was the analyses that were returned to him from his experts that started him thinking along revolutionary pathways.
4. He began writing in journals which he called his 'transformation notebooks.' This is where he worked out the specifics of his theory (which he called 'descent with modification. He never liked the name 'evolution' for his theory, because this term was already used for the development of an embryo from a fertilized egg, a process strictly controlled and predictable, which Darwin's theory says is not true for the history of life's diversity.
5. Darwin read Thomas Malthus's Essay on Populations. Malthus (1766-1834) was an economist who wrote that people always tend to over-reproduce, but that the resources they depend upon don't reproduce fast enough to continue to provide for the extended population. Thus, he said, if it weren't for things that control population size (like war, plague, and other pleasant things), there would never be enough resources to go around. Thus, in order for humans to continue to survive, some have to die.
6. Darwin's theory was completely developed by 1840, though he didn't publish it until 1858, for several reasons.
7. One reason he finally published in 1858 (and then published Origin of Species in 1859) was that he discovered that another man, Alfred Russell Wallace (1823-1913), had independently discovered and was about to publish exactly the same theory Darwin had been working on for 20 years.
B. What exactly is the theory of evolution? Darwin's theory has two novel concepts.
1. Darwin saw a clear pattern of common descent among the species of life. Examining the expert analyses of all of the specimens from the Beagle expedition, it was clear to Darwin that the various forms of life were all descended from one, or many from a very few, common ancestral species. Additional evidence since Darwin's time provides very powerful support for the idea of a single ancestral species for all life on Earth.
2. The mechanism Darwin figured out by which species could change from one thing into something different was natural selection. He'd spent virtually all of his life in the country, and was very aware of how radically artificial selection by plant and animal breeders could alter organisms. He saw environmental competition as providing the selective pressure in nature. Natural selection was partially derived from Malthus's ideas. Here's the reasoning:
a. Observation: All living things tend to over-reproduce.
b. Observation: Resources are finite.
c. Conclusion: There will be competition over resources, and not all members of any generation can survive.
d. Observation: There is variation in all species'a more profound statement than you might think
e. Observation: Variations in species can influence who can and who can't get enough resources to survive and reproduce.
f. Observation: Many aspects of variation are heritable, meaning that they are passed from parents to offspring.
g. Conclusion: Each new generation will tend to be more like the successful parents and less like the unsuccessful parents. Given enough time (meaning enough generations), these changes can produce brand new species, especially if the environmental pressure is strong.
IV. What is the evidence in support of the theory of evolution? This question could take weeks to answer. Observations of a wide variety of phenomena demonstrate evolutionary action and consequences. Here are some examples:
A. Artificial selection among plants and animals, as described above.
B. Adaptive radiation, which Darwin saw abundantly demonstrated among the species he observed and collected while on the Beagle. Adaptive radiation is the phenomenon in which a single species splinters into many daughter species. This phenomenon is very commonly observed among species living on off-shore islands like the Galapagos Islands to the west of South America.
C. The fossil record abundantly demonstrates evolutionary change. Transitional sequences show, for instance, the emergence of mammals from reptiles in the Permian Period, and the emergence of Cetaceans from terrestrial mammals in the early Cenozoic Era.
D. Biogeography (the distribution of species geographically) abundantly supports evolution, specifically island biogeography, which shows us that the resident species of islands are always derived from the species on the adjacent mainland.
E. Signatures of History are abundant in virtually all species. These are physical or behavioral features which clearly show that the modern species is descended from a species that lived a different kind of life cycle.
F. Molecular sequencing is an area of evidence that Darwin could never have predicted. Our modern biotechnology allows us to compare DNA and protein sequences among living organisms. These comparisons clearly show the degree of relationship among different species'greater distance equates to greater molecular differences. Molecular sequencing has provided a superb method of directly testing the predictions of evolutionary theory.
V. What is the modern status of Darwin's theory of evolution?
A. All of the basic concepts have remained sound, and have survived extensive challenge and testing. There was a great deal of evidence in support of Darwin's theory when he published it, and it has gained in acceptability and supportive evidence for nearly 150 years.
B. Do we agree with Darwin's original theory on all points? Of course not. 150 years of gathering and examining evidence has led to some greater understandings than were possible in Darwin's time.
1. One of the areas in which our available information is much greater is in the fossil record. We have many times more fossil evidence than Darwin did, and some things have become more clear with the greater abundance of physical evidence of extinct species. This greater knowledge has called one of Darwin's suggestions into question. Darwin felt that, to a large extent, new species arose from old species by the gradual accumulation of tiny changes, a pattern of change that has come to be called 'Darwinian gradualism.' If this were the typical pattern of new species production, we would predict that the fossil record would contain a very high percentage of transitional forms. In reality, though there are many transitional fossils, they don't occur in nearly the abundance predicted by gradualism. Niles Eldredge and Stephen Jay Gould proposed an alternative view, which has come to be called 'punctuated equilibrium.' They suggest that most species remain in stasis for long periods of time, and that when new species arise, they do so relatively rapidly (geologically speaking).
2. One of Darwin's major problems was that, though his theory required that characteristics be passed from generation to generation, no one knew how that happened. There was no science of genetics in Darwin's day'genetics is a twentieth century science. Darwin suggested a method of heredity which involved the production of what he called 'gemules' by various parts of the body, but it was incorrect. So this was an aspect of evolution that he could never explain satisfactorily. In the year 1900, the science of genetics was 'born.' Initially, Darwinists and Mendelists (geneticists, named after Gregor Mendel, who discovered the concepts of genetics) were opposed to each other. Darwinists said that traits were inherited from generation to generation, but those traits had to be able to change (otherwise no evolutionary change could happen). Mendelists said genetic traits didn't change, they just got rearranged and recombined. Growing understanding of mutation, which does change genes, allowed the two battling factions of biologists to come to an agreement. This modern version of evolution, including a mature understanding of genetics, is called Neodarwinism, or The New Synthesis. It is our modern theory of evolution'Darwin, only better.
VI. Some have greatly misused Darwin's ideas, particularly the concept of natural selection, to support their own notions of how humans should interact with each other. The theory of evolution is a theory about how the diversity of living things has come to be; it has nothing to do with human social systems, and should never be applied in these ways.
A. Social Darwinism is not actually 'Darwinism' of any kind. Charles Darwin himself was not a social Darwinist. It is from social Darwinists that we get the phrase, 'survival of the fittest.' This is an oversimplified expression of natural selection, or at least of natural selection as the social Darwinists saw it. Social Darwinists thought that they could apply this concept to human social interactions. Thus, in human interactions, the guy who could win was clearly innately superior. In war, the victory would go to the superior nationality. In business, the guy who could come out on top, no matter the strategies used, was obviously superior to the one who ended up on the bottom. Evolutionary theory has nothing to do with this sort of thing; this is just human politics and human philosophy at work, and trying to give itself legitimacy by dressing up in the trappings of science.
B. Eugenics was a specific expression of early twentieth century social Darwinism. The word literally translates as 'good origins.' The eugenics movement was based upon the idea that, as scientists finally understood both genetics and evolution, they could take control of the evolutionary future of the human species. Unfortunately, actual understanding of the complexities and realities of genetics and evolution was not really available. It still isn't available today. The primary concern of most eugenicists was the 'problem' of 'feeblemindedness,' which was felt to be at the root of all kinds of social ills. Terms like 'feebleminded,' 'moron,' and 'idiot' were not originally used as insults. They were technical terms given to particular ranges on the I.Q. 'bell curve.' The feebleminded were those in the segment one standard deviation below 100'those who were of 'substandard' I.Q., but who could function in the world'hold down jobs, marry, have children. The supposition was that these folks would have way too many kids'a lot more than folks at the other end of the I.Q. scale, and would thus drag the whole human species down with their 'stupid' genes. So the eugenicists advocated doing something about limiting the reproduction going on in this segment of the population. In the U.S. the preferred approach was sterilization.
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